Staff Publications

Listed below are technical publications by current Arkansas Natural Heritage Commission (ANHC) staff members (names in bold). Downloadable copies (PDF) of many articles are available below.

Birds

Holimon, W.C., C.W. Benkman, and M.F. Willson. 1998. The importance of mature conifers to red crossbills in southeast Alaska. Forest Ecology and Management 102:167-172.

Benkman, C.W., W.C. Holimon, and J.W. Smith. 2001. The influence of a competitor on the geographic mosaic of coevolution between crossbills and lodgepole pine. Evolution 53:282-294.

Holimon, W.C. and D.A. James. 2003. Nesting willow flycatchers discovered at Harrison, Arkansas. Journal of the Arkansas Academy of Science 57:81-85.

Holimon, W.C. and W.G. Montague. 2003. Reciprocal translocation re-establishes breeding status of Mississippi Alluvial Plain population of red-cockaded woodpeckers in Arkansas. Journal of the Arkansas Academy of Science 57:197-198.

Holimon, W.C., R.H. Doster, D.A. James, M.L. Mlodinow, J.C. Neal, and W.M. Shepherd. 2004. First documentation that Henslow's sparrow regularly occurs during the breeding and wintering seasons in Arkansas. Journal of the Arkansas Academy of Science 58:111-116.

Holimon, W.C., C.T. Witsell, W.H. Baltosser, and C.W. Rideout. 2008. Density and habitat associations of Henslow's Sparrows wintering in saline soil barrens in southern Arkansas. Journal of Field Ornithology 79(4):364-370.

Holimon, W.C., J.A. Akin, W.H. Baltosser, C.W. Rideout, and C.T. Witsell. 2012. Structure and composition of grassland habitats used by wintering Smith’s Longspurs: the importance of native grasses. Journal of Field Ornithology 83:351-361.

Fish and Aquatics

Bowman, D.W.,1993. Black bass in Beaver reservoir and its tributaries: distribution and abundance in relation to water quality. M.S. Thesis. University of Arkansas

Bowman, D.W., et al.1994. Angler use, success, and characteristics on Greers Ferry Tailwater, Arkansas, with implications to management. Proc. Annu. Conf. Southeast. Assoc. Fish and Wildl. Agencies, 48:499-511.

D. Bowman.1995. A new direction for trout management in Arkansas: developing wild trout populations in spring creeks. Published Abstract. Proceedings of the East Coast Trout Management Workshop II. May 31-June 2, Penn State University, University Park, Pennsylvania. American Fisheries Society Northeastern and Southern Divisions.

Rider, L.L. and D. Bowman. 1995. Results of a 1993 Arkansas trout angler survey with application to management. Published Abstract. Proceedings of the East Coast Trout Management Workshop II. May 31-June 2, Penn State University, University Park, Pennsylvania. American Fisheries Society Northeastern and Southern Divisions.

Stark, J. and D. Bowman. 1995. Movement of resident trout in the Bull Shoals Tailwater in response to differing levels of dissolved oxygen and spawning. Published Abstract. Proceedings of the East Coast Trout Management Workshop II. May 31-June 2, Penn State University, University Park, Pennsylvania. American Fisheries Society Northeastern and Southern Divisions.

Johnson, J.E., M.G. Pardew, and D. W. Bowman. 1996. Use of a nursery pond to establish smallmouth bass in Beaver Reservoir, Arkansas. Proceedings of the Annual Conference of Southeastern Association of Fish and Wildlife Agencies. 50:122-130.

Hudy, M., D. Downey, and D. Bowman. 2000. Successful restoration of an acidified native Brook Trout stream through mitigation with limestone sand. North American Journal of Fisheries Management. 20:453-466.

Williams, J.S., D.W. Bowman, C.S. Todd, M. Bivin, R. Moore. 2004. Multispecies trout management on a small Ozark tailwater. Proc. Annu. Conf. Southeast Assoc. Fish and Wildl. Agencies. 58:1-11.

Thompson, B., T. Scott, D. Bowman. Manuscript 2014. Using chemical fertilization as reservoir management technique to enhance biological productivity without degrading water quality and recreational use.

Knierim, K.J., Hays, P.D., and Bowman, D., 2015. Quantifying the variability in Escherichia coli (E. coli) throughout storm events at a karst spring in northwestern Arkansas, United States: Environmental Earth Sciences, v. 74, p. 4607-4623, doi: 10.1007/s12665-015-4416-5.

Mammals 

Witsell, C.T., G.A. Heidt, P.L. Dozhier, T. Frothingham, and M. Lynn. 1999. Recent documentation of mountain lion (Puma concolor) in Arkansas. Journal of the Arkansas Academy of Science 53:157-158.

Natural Communities

Foti, T.L. 1970. Correlation between selected soil and vegetation factors, Sylamore District, Ozark National Forest. U. of A. unpbl. M.S. Thesis.

Foti, T.L. 1975. The natural divisions of Arkansas. In Arkansas Natural Area Plan, Ark. Dept. of Planning, Little Rock, pp. 11-34.

Foti, T.L. 1976. Arkansas, its land and people. Arkansas Department of Education, Little Rock, 71p.

Foti, T.L. 1978. The Natural Divisions of Arkansas, a Classroom Guide. Arkansas Ecology Center, Little Rock, 70p.

Irving, R.S., S. Brenholts and T.L. Foti. 1980. Composition and net primary production of native prairies in eastern Arkansas. Am. Midl. Nat. 103(2): 298-309.

Foti, T.L. 1984. Environmental considerations of the prehistoric river valley. In P. C. Sherrod, Motifs of ancient man, pp. 130-135. UALR, Little Rock.

Foti, T.L. 1989. Blackland prairies of southwestern Arkansas. Proceedings of the Arkansas Academy of Science 43:23-28.

Foti, T.L. 1990. The vegetation of Saratoga Landing blackland prairie. Proceedings of the Arkansas Academy of Science 44:40-43.

Foti, T.L. and S.M. Glenn. 1992. The Ouachita Mountain landscape at the time of settlement. In: Hedrick, L. and D. Henderson, eds. Proceedings of the conference, Restoration of old growth forests in the Interior Highlands of Arkansas and Oklahoma. Ouachita National Forest, Winrock International Institute for Agricultural Development: 49-65.

Foti, T.L. and G. Hansen. 1992. Arkansas and the Land. 1992. U of A Press, Fayetteville. 113 pp.

Foti, T.L. 1993. The river's gifts and curses. In: J. Whayne and W.B. Gatewood eds. The Arkansas Delta, Land of Paradox. U of A Press, Fayetteville. pp. 30-52.

Foti, T.L. and M.S. Devall. 1994. Herbaceous plant biodiversity of stands in the Ouachita and Ozark national forests. In: Baker, J.B. 1994. Proceedings of a symposium on ecosystem management research in the Ouachita Mountain: pretreatment conditions and preliminary findings. New Orleans, LA: U.S. Department of Agriculture, Forest Service, Southern Forest Experiment Station.

Foti, T.L. and J.M. Guldin. 1994. Multivariate analysis of the ground cover layer, shrub layer, midstory, and overstory. In: Baker, J.B. 1994. Proceedings of a symposium on ecosystem management research in the Ouachita Mountain: pretreatment conditions and preliminary findings. New Orleans, LA: U.S. Department of Agriculture, Forest Service, Southern Forest Experiment Station.

Foti, T.L., X. Li, M. Blaney and K. Smith. 1994. A classification system for the natural vegetation of Arkansas. Proc. Ark. Acad. Science 48:50-62.

Foti, T.L. and G.A. Bukenhofer. 1998. A description of the sections and subsections of the Interior Highlands of Arkansas and Oklahoma. Journal of the Arkansas Academy of Science 52:53-62.

Foti, T.L. and G.A. Bukenhofer. 1999. Chapter 1: Ecological units of the Highlands. In Ozark-Ouachita Highlands Assessment: terrestrial vegetation and wildlife. Report 5 of 5. Gen. Tech. Rep. SRS-35. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southern Research Station: 1-6.

Foti, T.L., L.L. Richards, J. Andre and J.E. Voeller. 1999. Chapter 2: Prehistoric and historic ecological changes. In Ozark-Ouachita Highlands Assessment: terrestrial vegetation and wildlife. Report 5 of 5. Gen. Tech. Rep. SRS-35. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southern Research Station: 1-6.

Hamel, P.B. and T.L. Foti., tech. eds. 2001. Bottomland hardwoods of the Mississippi Alluvial Valley: Characteristics and management of natural function, structure and composition. A Symposium at Fayetteville, AR October 28, 1995. Gen. Tech. Rep. SRS-42. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southern Research Station. 109 p.

Tingle, J.L., C.V. Klimas, and T.L. Foti. 2001. Application of General Land Office Survey notes to bottomland hardwood ecosystem management and restoration in the Lower Mississippi Alluvial Valley - an example from Desha County, Arkansas. In: Hamel, P.B. and T.L. Foti tech. Eds. Bottomland hardwoods of the Mississippi Alluvial Valley: characteristics and management of natural function, structure, and composition. 1995 October 28. Fayetteville, AR. Gen. Tech. Rep. SRS-42. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southern Research Station: 16-27.

Foti, T.L. 2001. Presettlement forests of the Black Swamp area, Cache River, Woodruff County, Arkansas from notes of the first land survey. In: Hamel, P.B. and T.L. Foti, eds. Bottomland hardwoods of the Mississippi Alluvial Valley: characteristics and management of natural function, structure, and composition. 1995 October 28. Fayetteville, AR. Gen. Tech. Rep. SRS-42. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southern Research Station: 7-15.

Foti, T.L., S. Simon, D. Zollner, and M. Hattenbach. 2003. Blackland prairie landscapes of southwestern Arkansas: historical perspective, present status and restoration potential. pp. 94-109 In Blackland Prairies of the Gulf Coastal Plain, Nature, Culture and Sustainability, E. Peacock and T. Schausecker, eds. University of Alabama Press Tuscaloosa and London. 348 pp.

D. Zollner, S. Simon and T.L. Foti. 2003. A plant community classification for Arkansas’s blackland prairie ecosystem. Pp. 110-145 In Blackland Prairies of the Gulf Coastal Plain, Nature, Culture and Sustainability, E. Peacock and T. Schausecker, eds. University of Alabama Press Tuscaloosa and London. 348 pp.

Woods, A.J., T.L. Foti, S.S. Chapman, J.M. Omernik, J.A. Wise, E.O. Murray, W.L. Prior, J.B. Pagan, J.A. Comstock, Jr., and M. Radford. 2004. Ecoregions of Arkansas (color poster with map, descriptive text, summary tables and photographs): Reston, Virginia, US Geological Survey (map scale 1:1,000,000).

Foti, T.L. 2004. Upland hardwood forests and related communities of the Arkansas Ozarks in the early 19th century. In: Spetich, A., ed. Upland oak ecology symposium: history, current conditions, and sustainability. Gen. Tech. Rep. SRS-73. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southern Research Station: 21-29.

Klimas, C.V., E.O. Murray, J. Pagan, H. Langston, and T.L. Foti. 2005. A regional guidebook for applying the hydrogeomorphic approach to assessing wetland functions of forested wetlands in the West Gulf Coastal Plain region of Arkansas. ERDC/EL TR-05-12. US Army Corps of Engineers, Engineering Research and Development Center, Vicksburg, MS 159 pp. plus appendices.

Klimas, C.V., E.O. Murray, H. Langston, T. Witsell, T. Foti, and R. Holbrook. 2006. A regional guidebook for conducting functional assessments of wetland and riparian forests in the Ouachita Mountains and Crowley’s Ridge regions of Arkansas. ERCS/EL TR-06-14, U.S. Army Corps of Engineers.

Klimas, C.V., E.O. Murray, H. Langston, J. Pagan, C.T. Witsell, and T.L. Foti. 2008. A regional guidebook for applying the hydrogeomorphic approach to assessing wetland functions of forested wetlands and riparian areas in the Ozark Mountains region of Arkansas. ERDC/EL TR-08-31. US Army Corps of Engineers, Engineering Research and Development Center, Vicksburg, MS 116 pp. plus appendices.

Klimas, C.V., E.O. Murray, T.L. Foti, J. Pagan, M. Williamson and H. Langston. 2009. An Ecosystem restoration model for the Mississippi Alluvial Valley based on geomorphology, soils, and hydrology. Wetlands Vol. 29, No. 2, June 2009, pp. 430-450.

Seifert, C.L., R.T. Cox, S.L. Forman, T.L. Foti, T.A. Wasklewicz, and A.T. McColgan. 2009. Relict nebkhas (pimple mounds) record prolonged late Holocene drought in the forested region of south-central United States. Quaternary Research 71: 329-339.

Lockhart, Brian Roy, James M. Guldin, and Thomas Foti. 2010. "Tree Species Composition and Structure in an Old Bottomland Hardwood Forest in South-Central Arkansas." CASTANEA 75(3): 315-29. September 2010.

Bragg, D.C., R. O’Neill, W. Holimon, J. Fox, G. Thornton, and R. Mangham. 2014. Moro Big Pine: Conservation and Collaboration in the Pine Flatwoods of Arkansas. Journal of Forestry 112:446-456.

Plants

Irving, R.L., S. Brenholts and T.L. Foti. 1980. Composition and net primary production of native prairies in eastern Arkansas. American Midland Naturalist 103(2):298-309.

Witsell, C.T. and W.M. Shepherd. 1998. Rediscovery of Marsilea vestita subsp. vestita in Pulaski County, Arkansas after 162 years. Journal of the Arkansas Academy of Science 52:142-143.

Peck, J.H., C.T. Witsell, and E. Hendrix. 2000. Dryopteris goldiana and its hybrid with D. celsa new to Arkansas. American Fern Journal 90:110-111.

McKenzie, P.M., T. Smith, and C.T. Witsell. 2003. Carex conoidea (Cyperaceae) new to Arkansas with notes on its occurrence in Arkansas and Missouri. Sida 20:1727-1730.

Pringle, J.S and C.T. Witsell. 2005. A new species of Sabatia (Gentianaceae) from Saline County, Arkansas. SIDA 21:1249-1262.

Witsell, C.T. and B. Baker. 2006. The vascular flora of the South Fork Native Plant Preserve, Van Buren County, Arkansas. Journal of the Arkansas Academy of Science 60:144-164.

McKenzie, P.M., C.T. Witsell, and J. Woolbright. 2006. Carex bicknellii (Cyperaceae) new to Arkansas. Sida 22:801-804.

Dahal, P., H. Liechty, B.A. Rupar, C. Fristoe, E. Heitzman. 2006. Hardwood vigor and survival following applications of imazapyr in mid-rotation pine plantations. Gen. Tech. Rep. SRS-92. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southern Research Station. pp. 171-173.

Witsell, C.T. 2007. Hypericum adpressum (Clusiaceae) new to Arkansas and the Ouachita Mountains, U.S.A. Journal of the Botanical Research Institute of Texas 1:713-716.

Rupar, B.A. 2008. Growth response of Loblolly Pine and Competing vegetation to midrotation herbicide and fertilizer application. M.S. Thesis, University of Arkansas at Monticello, 85 p.

Jones, R.L., C.T. Witsell, and G.L. Nesom. 2008. Distribution and taxonomy of Symphyotrichium sericeum and S. pratense (Asteraceae: Astereae). Journal of the Botanical Research Institute of Texas 2:731-739.

Simpson, J.D. Crank, C.T. Witsell, and J.H. Peck. 2008. Two exotic ferns, Dryopteris erythrosora and Marsilea quadrifolia, newly naturalized in Arkansas. American Fern Journal 98:111-112.

McKenzie, P.M., C.T. Witsell, L.R. Phillippe, C.S. Reid, M.A. Homoya, S.B. Rolfsmeier, C.A. Morse. 2009. Status assessment of Eleocharis wolfii (Cyperaceae) in the United States. Journal of the Botanical Research Institute of Texas 3(2):831-854.

Weakley, Alan S., Richard J. LeBlond, Bruce A. Sorrie, C. Theo Witsell, L. Dwayne Estes, Kanchi Gandhi, Katherine Gould Matthews, and Atsushi Ebihara. 2011. "New Combinations, Rank Changes, and Nomenclatural and Taxonomic Comments in the Vascular Flora of the Southestern United States." Journal of the Botanical Research Institute of Texas 5(2): 437-55.

MacRoberts, Barbara R., Michael H. MacRoberts, and C. Theo Witsell. 2011. "Small-Scale Vascular Plant Species Richness in Southwestern Arkansas Blackland Prairies." Journal of the Botanical Research Institute of Texas 5(2): 743-51.

Gentry, J.L., G.P. Johnson, B.T. Baker, C.T. Witsell, and J.D. Ogle, eds. 2013. Atlas of the Vascular Plants of Arkansas. University of Arkansas Herbarium, Fayetteville, Arkansas.